The authors have tried to raise interspecific crosses between B. trilocularis or B. tournefortii and several other Brassica species, and succeeded in obtaining true F_1 hybrids in the following three cross-combinations: B. tournefortii x B. campestris, B. tournefortii x B. trilocularis and B. trilocularis x B. campestris. B. tournefortii was highly cross-incompatible with Brassica species having a, b or c genome, and B. trilocularis was, in turn, highly cross-compatible with Brassica species having a genome. Meiotic chromosome associations obtained with those F_1 hybrids were (0-4)Ⅱ+ (20-12)Ⅰ in F_1 B. tournefortii x B. trilocularis, (0-5)Ⅱ- (20-10)Ⅰ in F_1 B. tournefortii x B. canzpestris and (9-10)Ⅱ+ (2-0)Ⅰ in F_1 B. trilocularis x B. campestris. The result of genome-analysis has made clear that B. tournefortii is composed of the genome T, being partially homologous with the genome a, and that the genome a' of B. trilocularis is taken as that distinctly different from the genome a, though rather slight in its degree.
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